Had this same problem on my 2550 last year. This all started when we ran it dry feeding cows in the cold.
Did all the same things you did to try to solve the problem - filter, pump, new lines, clamps - nothing seemed to help.
Eventually, by using pressure to watch flow out of the tank, found a lot of junk in the bottom of the fuel tank. It would drain just fine with gravity, but the pressure simulated suction by trying to increase flow out of the tank. Since it holds a lot of fuel, could not see the outlet with a flashlight unless I drained the tank completely. I then saw the crud that was causing the issue. I would have not believed that the tractor could have run prior to running it dry. But now, for sure, the crud was migrating under suction causing a vacuum on the pump and pulling air. Drained the tank twice to get it clean and filled with fresh fuel, then everything started working correctly.
Still don't know why going dry was the catalyst for the issue.
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Today's Featured Article - A Lifetime of Farm Machinery - by Joe Michaels. I am a mechanical engineer by profession, specializing in powerplant work. I worked as a machinist and engine erector, with time spent overseas. I have always had a love for machinery, and an appreciation for farming and farm machinery. I was born and raised in Brooklyn, New York. Not a place one would associate with farms or farm machinery. I credit my parents for instilling a lot of good values, a respect for learning, a knowledge of various skills and a little knowledge of farming in me, amo
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